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Naturalist, Smithsonian African Expedition 




(Publication 2006) 



February 28, 1911 


Naturalist, Smithsonian African Expedition* 

Further study of the mammals collected by the Smithsonian 
African Expedition has made necessary the description of twenty- 
one additional new forms which are dealt with in the present paper. 
The bulk of these new forms are from Rhino Camp, in the Lado 
Enclave. The specimens in the collection are the first which have 
been obtained in this unknown region. The Lado Enclave com- 
prises the Nile watershed lying west of the Upper Nile (Bahr-el- 
Jebel). The precise geographical position of Rhino Camp is lati- 
tude 2° 55' north, on the west bank of the Nile some fifteen miles 
north of the station of Wadelai. 

This paper is the thirteenth dealing with the results of the expe- 
dition. Incidentally a new Lophuromys from West Africa is here 


Nimule Tree Squirrel 

Type from Uma (50 miles north of Nimule), Uganda; adult 
male, number 164828, U. S. Nat. Mus. ; collected by Kermit Roose - 
velt, February 11, 1910; Loring number, 8991. 

Characters . — Size of lateris , but coloration distinctively buffy, the 
feet and under parts pure buffy yellow. 

Coloration . — General coloration of upper parts mixed buffy, gray- 
ish, and black ; individual hairs annulated, basally black, then ochra- 
ceous, followed by a broad band of black and then by a narrower 
subterminal band of grayish buff and terminated by a black tip. 
Eyes with a broad whitish eye-rmg, cheeks and feet buffy. Tail 
annulated with buffy and black bands of equal width. Under parts 
and inside of limbs buffy ; under side of tail basally, and anal region, 
tawny rufous in marked contrast to rest of under parts. 

Measurements . — Head and body, 202 mm.; tail, 204; hind foot, 
44. Skull: condylo-incisive length, 41.5; basilar length, 35; zygo- 
matic breadth, 27.5; interorbital v T idth, 13.5; length of nasals, 14; 
upper toothrow, 9; condylo-incisive length of mandible, 30.5. 

Smithsonian Miscellaneous Collections, Vol. 56, No. 17 


This form of the widespread multicolor group is decidedly more 
buffy than its Lado ally, later is, approaching more the color of 
elegans of Mr. Elgon. It is, however, distinctly smaller than this 
form and somewhat yellower. 

The type is the only specimen in the collection. This was secured 
by Kermit Roosevelt in a clump of bamboo near the village of Uma. 

PARAXERUS KAHARI, new species 

Meru Forest Squirrel 

Type from Meru Boma, northeast of Mt. Kenia, British East 
Africa; adult female, number 164203, U. S. Nat. Mus. ; collected 
by Edmund Heller, September 24, 1909; original number, 1174. 

Characters. — Like ganana, but tail without rusty median stripe; 
skull larger and proportionately broader. 

Coloration . — General dorsal coloration buffy olive, darkest me- 
dially, becoming lighter and more buffy on the sides; individual 
hairs annulated by a narrow subterminal band of buff; overlaid by 
longer wholly black hairs ; dark area of back bordered by an obso- 
lete lighter stripe. Eye with a broad buffy ring. Cheeks and lower 
sides olive buff. Feet above ochraceous. Ears like dorsum. Tail 
mixed blackish and buffy, without marked darker cross-bands, the 
border light buffy ; below more yellowish buffy. Under parts cream 
buff, the hair on median line buffy to the roots. 

Measurements. — Head and body, 175 mm.; tail, 172; hind foot, 38. 
Skull: condylo-incisive length, 36.5; basilar length, 31.5; zygomatic 
breadth, 24.5; interorbital width, 11.5; length of upper toothrow, 
7.3 ; condylo-incisive length of mandible, 27.5. 

This race is closely allied to ganana of Jubaland. Specimens 
from the northern Guaso Nyiro River are exactly like the type in 
body coloration, but are somewhat smaller skulled. In coloration 
of upper parts this race is similar to ochraceus from Mombasa, but 
the coast species is at once distinguishable by its bright ochraceous 
under parts. 

Among the Wameru tribe this squirrel is known as kahari. 


Masked Dormouse 

Type from Rhino Camp, Lado Enclave; adult female, number 
164827, U. S. Nat. Mus.; collected by J. Alden Loring, January n, 
1910; original number, 8644. 


Characters. — Dorsal coloration buffy, as in lorraineus, but size 
much less. 

Coloration.— General dorsal colors buffy, overlaid by sepia, the 
buffy purer on sides and head and sharply defined against the gray 
of the under parts. Eye surrounded by a narrow black ring, which 
is continuous with a black streak to the tip of the snout. Cheeks 
creamy buff to level of eyes, where some fulvous separates it from 
the darker color of the upper parts; this light creamy color ascends 
behind ears and appears above them as a whitish tuft above each 
ear. Under parts grayish, washed with pale primrose yellow. 
Fore feet whitish, hind limbs broccoli brown like the tail; the toes 
and sides of the feet white. Tail above broccoli brown, the hairs 
lightening at tip and forming a light border; under surface lighter 
grayish or brown, but darkening toward tip, where it is the color 
of the upper surface. 

Measurements. — Head and body, 82 mm.; tail, 63; hind foot, 14; 
ear, 10. Skull: condylo-nasal length, 23; zygomatic breadth, 13.5; 
length of nasals, 8.5 ; interorbital width, 4.3 ; length of upper tooth- 
row, 3. 

Skull smaller than lorraineus , with shorter snout and smaller 
teeth. This species resembles lorraineus most closely in color and 
size. From smithii , which is about the same size, it differs widely 
in color, being buffy instead of grayish, with a distinct black eye- 

There is but a single specimen in the collection. 


Nii,e Biack-fronted Tree Mouse 

Type from Rhino Camp, Lado Enclave ; adult female, number 
164815, U. S. Nat. Mus.; collected by J. Alden Coring, January 18, 
1910; original number, 8783. 

Characters. — Like nigrifrons , but lighter, the under parts pearl 
gray; feet whitish and the upper parts cinnamon fawn. 

Coloration. — Color of upper parts cinnamon fawn, changing on 
lower sides and head to tawny buff, but not sharply defined against 
the pearl gray of the under parts ; a wide black median dorsal line 
from shoulders to root of tail, widest anteriorly and narrowing 
gradually posteriorly ; a median black spot on forehead midway 
between eye and ear. Snout lighter than rest of head, pure buffy 
yellow; base of whiskers and area between eye and tip of snout 



blackish. A few white hairs at anterior base of ears and a larger 
white patch immediately below ear. Ears dark seal brown with a 
few silvery hairs on inside. Eeet whitish. Lower parts pearl gray, 
the hair basally slate gray except on chin, throat, and under lip, 
which are white; median line of belly buffy. Tail dusky brown all 
around, essentially naked. 

Measurements . — Head and body, 68 mm. ; tail, y8 ; hind foot, 17-5* 
Skull: greatest length, 21; condylo-incisive length, 18.5; zygomatic 
breadth, 11.8; nasals, 7.5; diastema, 5; length of upper molar series, 
3.1 ; condylo-incisive length of mandible, 12. 

This species is allied to nigrifrons , but is readily distinguishable 
by its lack of buffy under parts and whitish feet. The series of 
eight topotypes shows very little variation in the color of the under 


Lado Tree Mouse 

Type from Rhino Camp, Lado Enclave; adult male, number 
164816, U. S. Nat. Mus.; collected by J. Alden Loring, February 1, 
1910; original number, 8921. 

Characters — Size of jamesoni and coloration of ochropus , with 
short ears and small skull and feet. 

Coloration . — Upper parts tawny ochraceous, lighter on the sides, 
ochraceous where the color is well defined against the white of the 
under parts; head lighter than body, ears darker umber-brown; a 
wide black median stripe extending from the shoulders to the base 
of the tail. Fore feet and under parts pure white, the hair white 
to the roots. Tail bicolor, dusky brown above, white below. 

Measurements . — Head and body, 65 mm.; tail, 87; hind foot, 
16.5. Skull: greatest length, 20; basilar length, 14.5; condylo- 
incisive length, 17.2; zygomatic breadth, 10; nasals, 7.2; diastema, 
5 ; length of upper toothrow, 3.5 ; condylo-incisive length of mandi- 
ble, 1 1.5. 

Twelve topotypes are in the collection. This series shows great 
variation in the distinctness of the black median dorsal stripe. In 
seven of these it is well defined, but in the remaining five there are 
all degrees of obsoleteness down to specimens which show only a 
very faint trace. 

Dendromus jamesoni of the Transvaal seems to be a close ally of 
this species, the chief differences separating the two forms being 
the lesser zygomatic width of the skull and the smaller ear of the 


Nile species. This new form approaches ochropus of British East 
Africa closer than any other equatorial species. From this species 
it is readily separable by the white under parts and the bicolored 
tail, and also by slightly smaller skull and feet. 

Two other specimens, both with stripe obsolete, are in the collec- 
tion from Iloima and Gondokoro, Uganda. 

MUS MUSCULOIDES EMESI, new subspecies 

Uganda Pygmy Mouse 

Type from Kabula Muliro, Uganda; adult male, number 164819, 
U. S. Nat. Mus. ; collected by J. Alden Loring, December 27, 1909; 
original number, 8497. 

Characters. — Dorsal coloration similar to murillus; hair of under 
parts white to the roots ; body size less, with proportionately smaller 

Coloration. — Median dorsal area blackish, sparingly flecked with 
tawny; sides lighter and more tawny, the lower sides well defined 
against the white of the under parts by a narrow band of fulvous ; 
dorsal pelage composed of black hairs annulated with tawny and of 
scattered wholly black hairs. Limbs like sides, ears and dorsal sur- 
face of tail blackish and clothed by minute hairs. Feet and under 
parts white, the hair white to the roots. Lower side of tail whitish. 

Measurements. — Head and body, 71 mm.; tail, 51; foot, 13.5. 
Skull: greatest length, 20.1; condylo- incisive length, 18.5; nasal, 
7.9; zygomatic breadth, 10.5; palatal foramina, 4.4; upper molar 
series, 3.5. 

Skull similar to murillus and to the members of the musculoides 
group generally, having the masseter knob and anterior cusp of 
m 1 little pronounced. This is a small white-bellied Nile race of the 
widespread musculoides group. 

The series of nine topotypes shows considerable variation in dor- 
sal coloration, ranging from the dark color of the type to a rufous 
phase. A specimen taken at Hoima, Uganda, is also referable to 
this new form. 

MUS WAMiE, new species 

Long-snouted Pygmy Mouse 

Type from Kapiti Plains, British East Africa; adult female, 
number 161777, U. S. Nat. Mus.; collected by J. Alden Loring, 
May 6, 1909; original number, 6061. 


Characters. — Long snouted like sorellus , but snout longer, the 
nasal bones measuring 8.5 mm. in length; upper parts much lighter, 

Coloration. — Median dorsal region from eyes to base of tail ochra- 
ceous-buff, heavily lined with black; snout and sides of head and 
body from level of eyes purer ochraceous-buff, changing to fulvous 
on lower sides where this color meets the white of the under parts. 
Ears hair brown, an ill-defined white spot at their base. Tail nearly 
naked ; light brownish above, whitish below. Under parts, feet, 
and snout from level of nose, white ; the hair white to the roots. 

Measurements. — Head and body, 68 mm.; tail, 40; foot, 15. 
Skull: occipito-incisive (greatest) length, 20.5; condylo-incisive 
length, 20; length of nasals, 8.5; width of interorbital constriction, 
3.2; length of palatal foramina, 5.4; width of brain case, 9; length 
of upper molar series, 3.6. 

This species is closely allied to sorellus, with which it agrees in 
the peculiar elongate snout and straight projecting incisors, and 
also in general proportions of body. Sorellus is, however, only 
distantly related to grains , but shows close relationship to tenellus 
in the formation of the short, narrowed mesopterygoid region, 
straight incisors and short tail. The group to which this species 
belongs represents the extreme divergence of the African Leggadas 
from the Mus muscitlits group in skull characters. This differ- 
ence from the type species of Mus is most marked in the straight, 
projecting incisors, the elongate snout, the abbreviated, narrow, 
mesopterygoid fossa, and the minute coronoid process of the man- 
dible. In the two latter characters these long-snouted African Mus 
approach somewhat the skull condition exhibited by Acomys. 

This is rather a rare species, being represented by only one skin 
and one alcoholic specimen in the collection, which contains some 
hundred specimens of Leggadas from British East Africa. 

MUS TENELLUS ACHOLI, new subspecies 

Long-snouted Pygmy Mouse 

Type from Rhino Camp, Lado Enclave; adult male, number 
164817, 1 T . S. Nat. Mus.: collected by J. Alden Loring, January 13, 
1910; original number, 8671. 

Characters. — Coloration of tenellus , but size much larger, with 
large skull and proportionately short tail. 

Coloration. — Median dorsal area mixed blackish and fulvous, the 



snout and posterior rump more fulvous; sides pure fulvous, fairly 
well defined against the dark dorsal area and the white of the under 
parts; limbs fulvous like the sides. Ears blackish, covered spar- 
ingly by minute grayish hairs. A conspicuous white subaural patch. 
Feet and under parts white, the hair white to the base. Tail hair 
brown above, whitish below, closely covered by short stiff hairs 
which hide the annuli. 

Measurements. — Head and body, 68 mm.; tail, 36; foot, 13. 
Skull: greatest length, 20.5; condylo-incisive length, 19; zygomatic 
breadth, 10; palatal length, 11; nasals, 8; length of upper molar 
series, 3.3; palatine foramina, 5. 

This is a larger race of the tenellus group of the Lower Sudan, 
of which it is the extreme in size. From snahelica of the Kiliman- 
jaro region this species differs widely in its much larger . size and 
lighter coloration. The tenellus group is welt defined by its long 
snout, projecting incisors, and short hairy tail. Three other speci- 
mens are in the collection from the type locality. 

MUS GRATUS SUNGAR7E, new subspecies 

Kenia Pygmy Mouse 

Type from West Kenia Forest Station, altitude 7,500 feet; old 
male, number 163487, U. S. Nat. Mus. ; collected by J. Alden Lor- 
ing, September 20, 1909 ; original number, 7425. 

Characters. — Like gratus, but larger and lighter colored, with 
the central dark area better defined. 

Coloration. — Median dorsal area from snout to base of tail mixed 
blackish and tawny in about equal parts ; sides lighter, purer tawny, 
changing somewhat abruptly from the dark dorsal region ; bounded 
on lower sides where the sides meet the white of the under parts by 
a fulvous line. Ears hair brown. Tail dark brown above, light 
brown below. Under parts and feet white, the hair white to the 

Measurements. — Head and body, 70 mm.; tail, 60; hind foot, 
14.5. Skull: occipito-nasal (greatest) length, 19.5; condylo-incisive 
length, 18.2; basilar length, 15.5; zygomatic breadth, 10; length of 
nasals, 7.2; length of palatal foramina, 4.9; length of upper tooth- 
row, 3.5. 

The material of this form in the collection is quite extensive. 
From the type locality there are six specimens, ten from Nyeri, 
seven from Waubugus, and eight from Naivasha. There is con- 


siderable variation due to age, the young usually having the under 
parts quite huffy with the hair basally plumbeous, the pure white 
under parts being acquired only in old adults. 

MUS BELLUS GONDOKORiE, new subspecies 

Gondokoro Pygmy Mouse 

Type from Gondokoro. Uganda; adult male, number 164820, U. S. 
Nat. Mus. ; collected by J. Alden Loring, February 25, 1910; orig- 
inal number, 9089. 

Characters . — Smaller and lighter colored than bellns , with the 
subaural white spot well defined. 

Coloration . — Median dorsal area from eyes to posterior rump 
hair brown, much lighter than the fulvous sides against which it is 
not well defined ; snout, area about eye, limbs, and sides of body 
bright fulvous, sharply contrasted with the white of the under parts. 
Ears hair brown like back. Under parts, tip of snout, and lips and 
feet, white ; the hair white to the roots. Tail brown above, lighter 
and whiter below. 

Measurements . — Head and body, 60 mm.; tail, 43; hind foot, 13. 
Skull: greatest length, 17.6; condylo-incisive length, 16; zygomatic 
breadth, 9.4; nasal, 63; length of upper toothrow, 3. 

The type is the only specimen in the collection. 

MUS BELLUS ENCLAVE new subspecies 

Enclave Pygmy Mouse 

Type from Rhino Camp, Lado Enclave ; adult male, number 
164818, U. S. Nat. Mus.; collected by J. Alden Loring, January 10, 
1910; original number, 8613. 

Characters . — Size of bclhts , but dorsal coloration much darker, 
the sides nearly as dark as the median region. 

Coloration. — L T pper parts blackish, the median region very spar- 
ingly flecked with tawny; sides more tawny, about equally mixed 
broccoli brown and tawny; snout dorsally blackish to tip, sides of 
head and snout tawny ochraceous, enclosing the eye on lower and 
posterior side only, the buffy eye-ring being broken above by the 
blackish suffusion of the head. Ears hair brown, with an indistinct 
white patch at their base. Tail dark brown above, lighter brownish 
below. Under parts from level of nostrils and feet white, the hair 
white to the base. 



Measurements. — Head and body, 64 mm.; tail, 46; hind foot, 13. 
Skull: occipito-nasal (greatest) length, 18; condylo-incisive length, 
16.5 ; length of nasals, 6.2 ; zygomatic breadth, 9.2 ; palatal foramina, 
4.3; length of upper toothrow, 3.1. 

This is a dark form of the widespread bellus . A series of thir- 
teen from Rhino Camp show very little variation in the dark upper 
parts and are readily distinguishable from the light sided bellus , 
with its sharply defined dark dorsal area. 


Lado Striped Grass Rat 

Type from Rhino Camp, Lado Enclave; adult female, number 
164825, U. S. Nat. Mus.; collected by J. Alden Loring. January 24, 
1910; original number, 8861. 

Characters. — Closely allied to massaicus, but feet smaller and 
skull narrower. 

Coloration. — General dorsal coloration bistre brown; the back 
marked by ten parallel lines of ochraceous spots, the central pair 
enclosing a median black dorsal line ; sides lighter and more grayish, 
the spots light buffy. Feet and ears ochraceous. Tail blackish 
above, buffy on sides and below. Under parts cream buff, the hair 
uniform to the roots. 

Measurements.— Uezd and body, 114 mm.; tail, 125; hind foot, 
23. Skull : condylo-incisive length, 25 ; zygomatic breadth, 23.5 ; 
nasals, 11x3; diastema, 7 ; length of upper toothrow, 5.2 ; length of 
bullae, 5 ; condylo-incisive length of mandible, 18. 

There are eleven topotypes in the collection. In color these range 
from the grayish color of massaicus to a buffy tint similar to ardeus. 
A series from Uganda, taken along the route from Kampala to 
Butiaba, agree in the small size of their feet and in color with the 
types series. One specimen from Gondokoro has the spots so 
coalesced as to form lines as in barbarus , from which it is not dis- 
tinguished except in the flatter skull and darker olivaceous body 


WhiTE-bElliEd Grass Rat 

Type from Rhino Camp, Lado Enclave; number 164826, U. S. 
Nat. Mus.; collected by J. Alden Loring. January 16, 1910; original 
number, 8762. 



voiv. 56 

Characters 'Pail long, length equaling head and body or greater; 
nasal bones slightly concave along lateral edges; upper parts tawny 
ocliraccous, lower parts white. 

Coloration . — Upper parts tawny ocliraccous, sparingly lined by 
black; purer ocliraccous on sides, the color extending low down on 
sides of belly, but not sharply defined against the* white under parts; 
pelage made up of narrowly seal brown tipped hairs with a broad 
baud of ocliraccous followed by slate to the base, the subterminal 
ocliraccous band giving the pelage its color character; interspersed 
with these annulated hairs are longer black ones, which give the 
median dorsal area a lined effect. Eyes with an ill-defined lmfTy 
ring. Ears chiefly ocliraccous, with a few black hairs about the 
base and on the inner side. Feet ocliraccous. Tail with a broad 
dorsal band of black, the sides huffy and the lower surface whitish. 
Under parts white, the hair white almost to the roots, the base ashy 
gray; on chin, lips, throat, and thighs the hair is white to the roots. 

Measurements . — Head and body, 140 mm.; tail, 150; hind foot, 
29. Skull: condylo-incisivc length, 31.5 ; greatest length, 35.5; 
zygomatic breadth, 17; interorbital constriction, 5; nasals, 12.5x4; 
length of upper row, 6.5; diastema, 8.2; length of palatine foram- 
ina, 5.5; length of bnlke, 5.6; condylo-incisivc length of mandible, 22. 

This species occurs at its type locality with rnbcsccns , both being 
forms of the abyssiniens group. Roth forms occur about Gondo- 
koro also. The color differences from the dark-bellied, shorter- 
tailed rubcscetis are marked, but the skull differences consist of only 
average characters, such as the slight concavity to the edges of the 
nasal hones and the lack of heading to the anterior part of the in- 
tcrorbital region. 

The type locality is represented by a series of twenty- four skins 
in which are specimens of all ages, but the variation in color is very 
little. The relationship between length of head and body and the 
length of the tail in this scries shows the following proportion : 
tail longer than head and body, 8 specimens; tail equal to head and 
body, 6 specimens; tail slightly shorter than head and body, 10 


Lado IIarsii -furred Mouse 

l'vpc from Rhino Camp, l v ado Enclave; adult male, number 
161823, S. Nat. Alus.; collected by J. Aldcn Eoring, January 24, 
1910; original number, 8853. 



Characters — Size large, skull Imig-snoutcd ; under parts bright 
orangc-ru fous. 

Coloration. — lJp]>er parts uniform seal brown, becoming lighter 
only on lower sides where the coloration merges gradually from 
fourth limber (o the cinnamon rufous of the belly. Feet lighter than 
back, vaudyke brown. Tail quite blackish above and on the sides, 
the median under surface becoming lighter. Under fur of upper 
parts chestnut brown. Under parts from chin to base of tail bright 
cinnamon rufous, the color uniform to the roots of the hair. 

Measurements . — Head and body, 136 mm.; tail, 7 2; hind foot, 
22.5. Skull ; condylo incisive length, 31 ; basilar length, 2 7; zygo- 
matic breadth, 15.5; inlerorbital width, 0.7; length of nasals, 14; 
length of upper tooth row, 5; condylo-iucisivc length of mandible, 

22 -s- 

Compared to imsorgei of the Kavirondo country of British East 
Africa, this form has the under parts much brighter red, the in- 
tensity of the red persisting through old age. Sikapusi, which has 
the same bright under parts, is a smaller race with a decidedly 
broader skull, with shorter, more convex nasal bones. 


Type from Kfulcu, lUda country, Kamcrun ; adult male, number 
125436, U. S. Nat. Mus.; collected by (). U. Bates, August 4, iQ° 3 > 
original number, 26. 

Characters. Tail essentially naked, the annulations conspicuous; 
hair very harsh; skull with small flattened hnllrc and shortened brain 
case, the parietal bones short. 

Coloration . — Upper parts uniform seal brown, the hair uniform 
in color to the roots on median line, but lightening on sides. Feet 
Vandyke brown. Under parts chiefly cinnamon rufous, the chest 
orange rufous. Tail quite naked, with distinct annulations, uni- 
form deep seal brown in color. 

Measurements . — Total length, 166 inm. ; tail (skin), 60; foot, 
18.5. Skull: condylo-incisivc length, 25; basilar length, 21; zygo- 
matic width, 13.5 (approximate) ; interorhital width, 6.5; length of 
nasals, ti.6; length of upper toothrow, 4.4; condylo-incisivc length 
of mandible, 17.3. 

'Phis is a small species of Cophnromys and differs from the other 
described species in its very much harsher fur and essentially naked 



VOL. 56 

URANOMYS UGANME, new species 

Uganda Big-toothed Mouse 

Type from Kikonda, Uganda; adult male, number 164822, U. S. 
Nat. Mus. ; collected by J. Alden Loring, January 2, 1910; original 
number, 8570. 

Specific characters. — Most closely related to ruddi of the Mount 
Elgon District of British East Africa. Differs decidedly in the 
shorter and narrower skull and the shorter nasal bones. Size 
smaller and coloration of under parts without any pinkish wash. 

Coloration. — Median portion of head and body above sepia flecked 
minutely with light brown, changing gradually on sides of head and 
body to a fulvous flecked with sepia. Under parts light grayish 
with a buffy wash. Fore and hind limbs with the color of the 
sides, except the feet, which are white. Tail dusky brownish above, 
becoming somewhat lighter below. 

Measurements. — Head and body, 101 mm.; tail, 75; hind foot, 18; 
ear, 12. Skull: greatest length, 27; condylo-basal length, 25; zygo- 
matic breadth, 14.5; nasals, 7.5 x 3.5 ; upper toothrow, 4.5; palatal 
foramina, 7.5. 

The skull of this genus is very close to Acomys in structure, the 
palatal region especially being similar, but externally this species 
can be scarcely distinguished from Lophuromys aquilas, with which 
it is found associated. The collection contains but a single speci- 
men. The teeth in this individual are quite worn, so that the tuber- 
cular structure cannot be made out. Compared with skulls of equal 
age of Lophuromys and Acomys , the m 3 is found to lack the minute 
posterior median cusp which is found in both these genera. The 
structure of the floor of the skull closely resembles Acomys. The 
depressions marking the termination of the bony palate are, how- 
ever, much deeper and have prominent margins, the interpterygoid 
fossa is wider and has parallel sides and blunt bead-shaped ptery- 
goid processes. The mandible resembles more that of Lophuromys 
in the shape of its coronoid process, which is long and slender and 
very different from the minute projection which is found in Acomys. 
The condylar process is considerably larger than in either Lophu- 
romys or Acomys. The most marked difference, however, is the 
prominence given to the incisor capsules. In this genus the capsule 
forms a prominent shelf-like projection midway between the coro- 
noid and the condylar processes. The incisor condyle of Acomys and 



Lophuromys is scarcely perceptible on the surface of the posterior 
portion of the ramus. This genus has the external characters of 
Lophuromys with the palatal skull formation of : A corny s and dis- 
tinctive projecting incisors and marked mandibular characters. 

ACOMYS HYSTRELLA, new species 

Gray-backed Spiny Mouse 

Type from Nimule, Uganda; adult female, number 164821, U. S. 
Nat. Mus. ; collected by J. Alden Loring, February 4, 1910; original 
number, 8929. 

Characters. — Size large ; tail three-fourths head and body ; dorsal 
coloration drab gray with fulvous sides; nasal bones short. 

Coloration. — Median dorsal region from the snout to the base of 
the tail drab-gray, this color merging gradually into the fulvous 
of the sides; fulvous of cheeks, lower sides, and legs sharply defined 
against the white of the under parts. Spinous hair of sides annu- 
lated, the extreme tip seal brown, succeeded by a broad band of 
fulvous, which is followed by cinereous gray to the base ; this annu- 
lation is not perceptible in the darker spines of the back, the sub- 
terminal band of drab-gray being nearly as dark as the tips. Feet 
white. A small white spot below the ear. Ears smoky-brown 
clothed with a mixture of minute brownish and silvery hairs. Tail 
sharply bicolor; above lighter gray than the back, below whitish. 
Under parts, including upper lips and feet, white, the hair white to 
the roots. 

Measurements. — Head and body, in mm.; tail, 81 ; hind foot, 16; 
ear, 15. Skull: greatest length, 28.3; basilar length, 22; zygomatic 
breadth, 13.2; nasals, 10.5x3.5; interorbital constriction, 5; length 
of palatine foramina, 6.7; length of upper tooth row, 4.5. 

From the Somali species, mullah, this species differs in its smaller 
size, more fulvous sides, and shorter tail and nasal bones. The ful- 
vous sides also distinguish it from the grayish cahirinus of Egypt, 
from which it further differs by larger skull and acutely pointed 
mesopterygoid fossa and smaller bulke. 

DASYMYS ORTHOS, new species 

Nyanza Swamp Rat 

Type from Butiaba, Albert Nyanza, Uganda; adult male, number 
164824, U. S. Nat. Mus. : collected by J. Alden Loring, January x, 
1910; original number, 8601. 



VOL. 56 

Characters - — Antorbital plate with a straight anterior margin, 
with scarcely any concavity; bullae small, coloration brownish; size 

Coloration. — General dorsal coloration Prout’s brown, becoming 
grayer on the sides and merging gradually into the olive-yellow 
wash of the gray under parts. Dorsal pelage made up of long 
black hairs which overlay the nearest shorter ones, which are 
terminally seal brown, then broadly tawny and basally slate. Limbs 
light wood brown. Tail showing very little hair, the amiulation 
conspicuous, color broccoli brown. 

Measurements. — Head and body 160 mm.; tail, 156; hind foot, 
32. Skull: condylo-incisive length, 36.5; zygomatic breadth, 18.5; 
nasal, 13.5; palatal foramina, 8; length tympanic bulla?, 5.6; length 
upper toothrow, 7.3; diastema, 11.2; condylo-incisive length of 
mandible, 26. 

This species is a much browner species than medius , with larger 
body size. The skull differs decisively from a large series of 
helnkus from British East Africa in its straight-edged antorbital 
plate, which is scarcely more angulated at its upper edge than 
in Epimys alexandrinns . 

There is one other specimen from Butiaba in the collection, and 
a third from Rhino Camp. 

DASYMYS SAVANNUS, new species 

Kenia Swamp Rat 

Type from Fort Hall, British East Africa; adult female, number 
164471, U. S. Nat. Mus.; collected by J. Alden Loring, October 28, 
1909; original number, 8182. 

Characters. — Coloration umber-brown, skull with antorbital plate 
much angulated and produced, giving the antorbital vacuity great 
depth; size smaller than in helnkus; tail essentially naked. 

Coloration. — Upper parts umber, overlaid medially with black, 
the individual hairs tawny-ochraceous, with dark brown tips and 
slaty bases mixed with longer black hairs. Sides lighter, purer 
ochraceous, becoming more grayish where they meet the olive-gray 
wash of the lower parts. Feet pale brown. Ears hair brown, 
clothed with minute grayish hairs. Tail like the ears in color, but 
quite naked, with the annuli showing conspicuously. 

Measurements. — Head and body, 156 mm.; tail, 146; hind foot, 
29.5. Skull : condylo-incisive length, 37.5 ; zygomatic breadth, 19 ; 



nasal, 14; palatal foramina, 8.5; length of bullae, 6; upper tooth- 
row, 7.5; diastema, 12.5; condylo-incisive length of mandible, 28. 

This species is very much browner than hclukus, with a less hairy 
tail in old age, and smaller body size. The angle of the antorbital 
plate is much more pronounced than in any other specimen in the 
collection. Interorbital region narrow, the beads on margin much 
raised and leaving a deep median groove. 

A large series of specimens show the age differences in skulls to 
be great. The interorbital region changes from a flat, weakly 
beaded condition to a narrow, deeply concave affair with high beads. 
Bulke in youth high and rounded, becoming in age flat and broad, 
antorbital plate less concave in young. 


Kenia Forest Mungoose 

Type from west slope of Mt. Kenia, altitude 8,500 feet; adult 
male, number 164152, U. S. Nat. Mus. ; collected by J. Alden Lor- 
ing, October 12, 1909; original number, 7934. 

Characters. — Much darker and larger than sanguineus , with black 
predominating on the dorsal surface ; skull larger and wider zygo- 
ma tically. 

Coloration . — Median dorsal area from snout to tip of tail black- 
ish ; top of head and tip of tail wholly black; back and sides mixed 
tawny and black, the black predominating; feet like back, only the 
toes black; sides of face black like the crown. Under parts mixed 
tawny and dark brown, the tawny predominating. 

Measurements. — Head and body, 300 mm.; tail, 282; hind foot, 
59. Skull: condylo-basal length, 65; palatal length, 34; zygomatic 
breadth, 34.5 ; width of brain case, 26.5 ; greatest diameter of pm 4 , 8. 

The series of six topotypes shows considerable variation. The 
darkest one has the head feet and median line of the back wholly 
black. The lightest one is somewhat darker than normal specimens 
of sanguineus. This new race is nearest to proteus of Mt. Ruwen- 
zori, which lives in very similar environment. It has a longer tail 
and larger skull, and is much less blackish. 



VOL. 56 


East African Banded Mungoose 

Type from the Southern Guaso Nyiro River, Sotik District, Brit- 
ish East Africa; adult female, number 162132, U. S. Nat. Mus. ; 
collected by Edmund Heller, June 21, 1909; original number, 6396. 
J. Alden Loring. 

Characters . — Size and proportions as in macritras , but coloration 
much grayer, the rufous of the shoulders replaced by buffy gray 
and the feet seal brown. 

Coloration . — General dorsal coloration buffy grayish, lined every- 
where with blackish ; the posterior back and rump marked by eleven 
dark cross bands which are well defined only on back, being obso- 
lete on the sides. Separating the broad dark cross bands are nar- 
rower light bands of buffy, which on median line become ferrugi- 
nous. Shoulders and limbs uniform in color with the general body 
color, the feet and the muzzle dor sally from the level of the eyes 
dark seal brown. Tail like body color, the terminal one- fourth 
blackish. Under parts with an ochraceous wash, the hairs basally 
dark brown ; throat lighter buffy, the chin and upper lip bright 
tawny, in marked contrast to rest of under parts. 

Measurements . — Head and body, 380 mm.; tail, 260; hind foot, 92. 
Skull : condylo-basal length, 7 5.5 ; basal length, 71 ; zygomatic 
breadth, 42.5 ; palatal length, 39; greatest diameter of pm 4 , 7. 

This East African veldt race of the banded mungoose is a much 
grayer and lighter colored animal than macrurus of Ruwenzori and 
Uganda. The rich rufous suffusion of the shoulders and the black 
feet of the latter are wanting, the body color being quite uniform 
and the black replaced by brown. 

These mungooses are often met with on the grassy plains of the 
Sotik country, where they live in colonies in burrows on the open 
veldt. They, however, do not stop long in any locality, but move 
about in small packs of ten to twenty individuals, which take up a 
temporary abode in any nest of burrows which they find convenient. 
From our observations it was apparent that they do not remain more 
than a day or two in any one set of burrows.